The story begins at Kaminoyama High School, where five close friends—Aoi Miyamori, Ema Yasuhara, Midori Imai, Shizuka Sakaki, and Misa Toudou—discover their shared passion for anime and establish an animation club. After producing their first amateur anime and presenting it at the school's culture festival, the group pledges to build careers in the industry, with the goal of eventually working together to create their own mainstream series. Two and a half years later, Aoi and Ema have secured positions at the prestigious Musashino Animation studio. The others, however, struggle to obtain their ideal roles. Shizuka feels the pressure of not being acknowledged as a competent voice actress, Misa holds a stable but unfulfilling job designing 3D models for an automotive company, and Midori is a university student determined to realize her dream of becoming a story writer. These five girls will discover that the road to success is filled with detours, yet dreams can still be realized through persistence and a dash of unconventional creativity.