A girl named Eruruu discovers an injured man in the woods, and his entire background remains a mystery. Lacking any knowledge of his past or even his own name, he is taken into Eruruu's home and given the name Hakuoro by her grandmother and younger sister, Aruruu. Although the village residents possess large ears and tails, Hakuoro's distinct physical characteristic is quite different: he has neither ears nor a tail, but wears a mask that he cannot remove. Shortly after integrating into the villagers' lives, a revolution against the land's tyrannical emperor erupts, bringing the conflict to his new home. Hakuoro must do everything in his power to protect the people and the village he has come to love, all while unraveling the mysteries surrounding his past.