Wonderful Precure! takes place in Animal Town, a community where humans and animals coexist peacefully. Iroha Inukai, a second-year junior high school student, shares a close friendship with her dog, Komugi. During a walk, Iroha and Komugi come across a mysterious being called Gargal, who begins to wreak havoc in the city. To safeguard Iroha, Komugi unexpectedly transforms into the Pretty Cure known as Cure Wonderful.