Compelled to flee their home world of Cybertron because of a black hole threat, Optimus Prime gets ready to guide his Autobot fighters to Earth. Right as they depart, the Autobots are met by Vector Prime, who cautions them about the danger to the galaxy if the four Cyber Planet Keys are not collected in time. Meanwhile, Megatron has his own designs for the universe. During their quest, they journey to new planets populated by various Transformer races, as well as the Autobots who rule these worlds.