The narrative is set one year after the OVA. Due to the previous war, Kabucki-cho was devastated, prompting the owner to abandon his lingerie pub, "Pallion," and open "New Papillion" in Akihabara. During the conflict, Tsubomi, along with her colleagues Anne and Shizuku, lost their memories and started new lives in Akihabara. Meanwhile, the Susano, aliens invading Earth, appeared. They transformed a UMA (Unidentified Mysterious Animal) into a monster and caused it to raid Akihabara. Rama the cat enabled Tsubomi and her friends to recover their memories, allowing them to fight the monster as Lingerie Soldiers.