Tetsuro Hoshino is a young boy born into poverty on Earth, who desires nothing more than a machine body. He also seeks revenge against Count Mecha, an evil man who was given a machine body and now hunts humans for sport, for killing his mother. As a street urchin, Tetsuro has no means to purchase a machine body. However, he has heard that at the end of the Andromeda Galaxy, there exists a planet where such robot bodies are given away for free. All Tetsuro needs to do is obtain a pass for the Galaxy Express 999, the cosmic railway. One day, Tetsuro meets the mysterious Maetel, who promises to give him her unlimited use pass for the Express on the condition that she be his travelling companion. Tetsuro happily agrees, and embarks with Maetel on an adventure across the stars. When Tetsuro starts out, he is every bit the whiny, immature, selfish child. However, the adventures he goes on with Maetel, the people he meets, and the sights he witnesses cause Tetsuro to slowly but truly grow into a strong, young man. Tetsuro also grows closer and closer to Maetel, and although the nature of their relationship might not be called romantic, they form a very strong bond of love.
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