Jeanne is a girl who grew up in the country of Donremi on the Western Planet. She experiences heavenly visions of the birth of a "Star Messiah" who will save the world, which leads her to embark on a journey to the Eastern Planet with Leonardo da Vinci, "the one who observes the world." On that planet, they encounter the heretic of the Eastern Planet and "the greatest fool of the day," Oda Nobunaga. Oda has her disguise herself as a man and gives her the name Ranmaru to protect her from their kingdom's custom, where women are treated as a secondary class.
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